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Based on genome sequencing data, it is estimated that the class of insects originated on Earth about 480 million years ago, in the Ordovician, at about the same time terrestrial plants appeared. Insects evolved from a group of crustaceans. They were the first animals to develop flight, about 400 million years ago in the Devonian period.〔 The oldest definitive insect fossil, ''Rhyniognatha hirsti'', is estimated to be . Global climate conditions changed several times during the history of Earth, and along with it the diversity of insects. The Pterygotes (winged insects) underwent a major radiation in the Carboniferous (356 to 299 million years ago) while the Endopterygota (insects that go through different life stages with metamorphosis) underwent another major radiation in the Permian (299 to 252 million years ago). Most extant orders of insects developed during the Permian period. Many of the early groups became extinct during the mass extinction at the Permo-Triassic boundary, the largest extinction event in the history of the Earth, around . The survivors of this event evolved in the Triassic (252 to 201 million years ago) to what are essentially the modern insect orders that persist to this day. Most modern insect families appeared in the Jurassic (201 to 145 million years ago). In an important example of co-evolution, a number of highly successful insect groups — especially the Hymenoptera (wasps, bees and ants) and Lepidoptera (butterflies) as well as many types of Diptera (flies) and Coleoptera (beetles) — evolved in conjunction with flowering plants during the Cretaceous (145 to 66 million years ago). Many modern insect genera developed during the Cenozoic that began about 65 million years ago; insects from this period onwards frequently became preserved in amber, often in perfect condition. Such specimens are easily compared with modern species, and most of them are members of extant genera. ==Fossils== Insect fossils are not merely impressions, but also appear in many other forms; While wings are indeed a common insect fossil, they do not readily decay or digest, which is why birds and spiders typically leave the wings after devouring the rest of an insect. Terrestrial vertebrates are almost always preserved just as bony remains (or inorganic casts thereof), the original bone usually having been replaced by the mineral apatite. Occasionally, mummified or frozen vertebrates are found, but their age is usually no more than several thousand years. Fossils of insects, in contrast, are preserved as three-dimensional, permineralized, and charcoalified replicas; and as inclusions in amber and even within some minerals. There is also abundant fossil evidence for the behavior of extinct insects, including feeding damage on fossil vegetation and in wood, fecal pellets, and nests in fossil soils. Dinosaur behavior, by contrast, is recorded mostly as footprints and coprolites.〔 The common denominator among most deposits of fossil insects and terrestrial plants is the lake environment. Those insects that became preserved were either living in the fossil lake (autochthonous) or carried into it from surrounding habitats by winds, stream currents, or their own flight (allochthonous). Drowning and dying insects not eaten by fish and other predators settle to the bottom, where they may be preserved in the lake’s sediments, called lacustrine, under appropriate conditions. Even amber, or fossil resin from trees, requires a watery environment that is lacustrine or brackish in order to be preserved. Without protection in anoxic sediments, amber would gradually disintegrate; it is never found buried in fossil soils. Various factors contribute greatly to what kinds of insects become preserved and how well, if indeed at all, including lake depth, temperature, and alkalinity; type of sediments; whether the lake was surrounded by forest or vast and featureless salt pans; and if it was choked in anoxia or highly oxygenated. There are some major exceptions to the lacustrine theme of fossil insects, the most famous being the Late Jurassic limestones from Solnhofen and Eichstätt, Germany, which are marine. These deposits are famous for pterosaurs and the earliest bird, Archaeopteryx. The limestones were formed by a very fine mud of calcite that settled within stagnant, hypersaline bays isolated from inland seas. Most organisms in these limestones, including rare insects, were preserved intact, sometimes with feathers and outlines of soft wing membranes, indicating that there was very little decay. The insects, however, are like casts or molds, having relief but little detail. In some cases iron oxides precipitated around wing veins, revealing better detail.〔 There are many different ways insects can be fossilized and preserved, one which includes compressions and impressions, concretions, mineral replication, charcoalified (fusainized) remains, and their trace remains. Compressions and Impressions are the most extensive types of insect fossils, occurring in rocks from the Carboniferous to Recent. Impressions are like a cast or mold of a fossil insect, showing its form and even some relief, like pleating in the wings, but usually little or no color from the cuticle. Compressions preserve remains of the cuticle, so color distinguishes structure. In exceptional situations, microscopic features such as microtrichia on sclerites and wing membranes are even visible, but preservation of this scale also requires a matrix of exceptionally fine grain, such as in micritic muds and volcanic tuffs. Because arthropod sclerites are held together by membranes, which readily decompose, many fossil arthropods are known only by isolated sclerites. Far more desirable are complete fossils. Concretions are stones with a fossil at the core whose chemical composition differs from that of the surrounding matrix, usually formed as a result of mineral precipitation from decaying organisms. The most significant deposit consists of various localities of the Late Carboniferous Francis Creek Shale of the Carbondale Formation at Mazon Creek, Illinois, which are composed of shales and coal seams yielding oblong concretions. Within most concretions is a mold of an animal and sometimes a plant that is usually marine in origin. When an insect is partly or wholly replaced by minerals, usually completely articulated and with three-dimensional fidelity, is called Mineral replication. This is also called petrifaction, as in “petrified” wood. Insects preserved this way are often, but not always, preserved as concretions, or within nodules of minerals that formed around the insect as its nucleus. Such deposits generally form where the sediments and water are laden with minerals, and where there is also quick mineralization of the carcass by coats of bacteria. 抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』 ■ウィキペディアで「Evolution of insects」の詳細全文を読む スポンサード リンク
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